A Short History of California Dragonflies aka Odonata
As recalled by Kathy Biggs (with help from Tim Manolis)
November 2006; updated February 2008
The first in-depth assessment of California's Odonata fauna was by Clarence Kennedy in 1914-1917. When I give programs, I often refer to him as the "Grandfather of California Dragonflies." I link to a few of his beautiful original drawings on my California Dragonflies Website. His Notes on the Life History and Ecology of the Dragonflies (Odonata) of Central California and Nevada (1917, Proceedings of the U. S. National Museum, Vol. 52-No.2192), which summarizes most of this field work, remains well worth re-reading today, primarily for the wealth of information on identification, behavior, and life histories of the state's dragonflies.
In 1956 Smith and Pritchard's "Odonata" chapter in Aquatic Insects of California provided keys to dragonflies down to species, but included other western species not known to occur in California. This was Tim Manolis's and my original guide when we started looking at Odes. It includes drawings from Kennedy. I link to a few of these beautiful drawings on my California Dragonflies Website: http://southwestdragonflies.net/caphotos/SNAKESKN.JPG and http://southwestdragonflies.net/caphotos/SNAKSKN2.JPG and http://southwestdragonflies.net/caphotos/BRIMSTN.GIF and http://southwestdragonflies.net/caphotos/OLIVACE.GIF.
Then the first Odonata species list for California was compiled by Dennis Paulson
(1975 photo) and Rosser
Garrison and published in April 1977
in The Pan-Pacific Entomologist, A List and New Distributional Records of Pacific Coast Odonata: 53:147-160.
It showed 102 species. At the 2003 DSA meeting in California, CalOdes Members awarded Dennis and Rosser special DSA
Pins proclaiming them to be the "Fathers of California Dragonflies." Dennis is now retired and a member of our CalOdes
group and lives in Washington. Rosser is in Sacramento working for the California Department of Food and Agriculture.
Both Dennis and Rosser are world-renowned odonatologists and have been very generous with their time and expertise in
helping us learn about California's dragonflies.
I started looking at dragonflies in 1996 when we built our backyard pond in Sebastopol and they appeared there. Soon I realized that I had more species than shown in Powell & Hogue's California Insects, so I started doing research. There were no identification books in the library or at the bookstores, so I started using the Internet (which was `new' back then!).
Also in 1996, Ron Lyons posted A list of California Dragonfly Species on a website, largely based on the list that Dennis and Rosser had compiled in 1977, thus making the list available to anyone on the Internet. Ron, now a CalOdes member even though he has moved to Oregon, has left an archival version of this website on the Internet:
I first found Ron's list and then another website with a list of species and links to photos. The second website was Blair Nikula's Dragonflies and Damselflies of Cape Cod.. I printed out Ron Lyon's list and then would print out a photo from Blair's website if I found a match, and then take this binder booklet into the field with me.
In August of 1996 the Dragonfly Society of the Americas (DSA) voted on Common Names for dragonflies. Dennis Paulson and Sid Dunkle were instrumental in forming this list of common names for North American Dragonflies.
Two new dragonfly resources became available later in 1996: Forrest Mitchell developed a technique for scanning
live dragonflies and put the scans of his Texas Dragonflies on his
Digital Dragonflies Website.
Dennis Paulson published the list North American Odonata,
The Odonata of North America with Common (English) Names on the web at his
Dragonfly Biodiversity website with Common Names of North American
Dragonflies. The updated website is available at the link above. He next began putting Scans of
Washington's Odonata on the web. He also included a Field Key to
Washington Dragonflies which worked fairly well here in California. I was therefore able to expand the number of
photos of California species I could print out in my binder, and Dennis answered my emails and helped me with
identification skills. I've always considered Dennis to be my personal mentor.
Ron Lyon's original California Dragonfly list was by scientific name only (it was posted before the Common Names were assigned). To thank Ron for all the help his website had been, I asked if I could add the newly assigned common names to it. He said yes and I did. Then I found that I wanted to add more to it, such as links and descriptions. So, with his permission I started a duplicate site in 1997: California Dragonflies and Damselflies aka California Odonata. I kept adding to the website as I discovered more information and made the decision to share as much of this information as I could so others wouldn't have to duplicate my efforts.
The website and my binder booklet kept growing. In late 1997 I received an email from another Californian who was interested in Dragonflies: Tim Manolis. He and I made a decision to go out the next year and see how many dragonfly species we could find. We did this starting in 1998. Tim read an article in the newspaper about a student at UCD who was studying dragonflies: Andy Rehn. We made an appointment to meet with him at the Bohart Insect Collection at UCD and we were given information on where Andy had found species and how to identify them.
We were now aware of how little information there was on the distribution of Dragonflies in California, and at Tim's and my request in April of 1998, Dennis Paulson sent a list with species by county to us on an Excel chart. There were 103 species on it with the Kiowa Dancer, Argia immunda, as the additional species added since his 1977 paper (actually collected in 1955!). I have kept an archival copy of this spreadsheet. It had only "zeros" and "ones" on it, where the "zeros" represented literature records, and "ones" represented specimens examined by Dennis in his own collection, and the collections of the California Academy of Sciences (CAS), the California Insect Survey collection (CIS), and the Los Angeles County Museum of Natural History (LACNNH).
Tim, Andy, and I quickly added 5 new numbers to the CA_OD_counties excel chart: 2 - UCD/Bohart Museum collection examined by A. Rehn &/or T. Manolis; 3 - Cal. State Univ., Chico by T. Manolis &/or A. Rehn; 4 - Cal. State Univ., Sacramento by T. Manolis &/or A. Rehn; 5 - CAS - CA Acad. of Sciences - Manolis, Biggs, Rehn; and our own research: 6 - collection of Tim Manolis; and 7 - collection of Kathy Biggs. Other folks soon began bringing their specimens and photo collections to us for review, including Rich Stallcup, Jeff Cole, Alvaro Jaramillo, Greg Kareofelas, Ron LeValley, David Wyatt, and others. In 1998 I started a sightings site to record our findings.
It was also in 1998 that Tim Manolis and Andy Rehn added three new species for
the state -
Belted Whiteface, Leucorrhinia proxima; Canada Darner, Aeshna canadensis; and Autumn Meadowhawk,
Sympetrum vicinum -- the first two collected at Willow Lake in Tehama County, the latter species collected in 1980
(found in the Bohart Museum at UC Davis in a drawer of previously unidentified specimens) by Andy Rehn, and found again in
1998 at Willow Lake. State list at 106 species.
My girlfriends had found I was always either out chasing dragonflies or working on the website. They teased me that I should write a field guide. In July of 1999 I took the California dragonflies binder along with me on the Sonoma County Butterfly count. Needless to say, I counted butterflies and dragonflies. Also on this trip was Doug Vaughan. He asked if I could give/sell him a copy of this information. That gave me the nerve to tell him that, although I couldn't because these weren't images I had permission to distribute, I had started writing my own field guide. Doug's wife, Doris Kretschmer, was editor for the Natural History Guides series at University of California Press. A call from her took me to the UCPress offices to meet her. She said UCPress was interested in publishing a dragonfly guide and asked if anyone I knew or I would be interested in writing one. I knew what I wanted to write, a simple beginner's guide, and they were interested in more than that. So I told them about Tim Manolis.
It was also in 1999 that Jeff Cole discovered that the specimens we had all
collected that spring at Dos Palmas (Kathy and Dave Biggs, Jeff Cole, Rosser and Jo Garrison and family, Tim and Annette
Manolis) were not Desert Forktails, Ischnura barberi, as we had presumed, but Rambur's forktails, Ischnura ramburii!! The species, originally removed by Dennis and Rosser in 1977, was now added back to state list. State list at 107
species.
In May of 2000, my Common Dragonflies of California, A Beginner's Pocket Guide
was published by Azalea Creek Publishing (Dave and Kathy Biggs) and I began giving
programs about dragonflies. Later that year, Sid Dunkle's Dragonflies through Binoculars was published by
Oxford Press.
Also in 2000 we discovered that a specimen of the Riffle Darner, Oplonaeschna armata had been collected in 1964 in California. It was found in the Florida State Collection of Arthropods and was discovered as part of Nick Donnelly's research for the USA spot-map project. We were quite busy that year helping with that project by sharing our spreadsheet information with Nick and visa versa. Nick's maps became the basis of the information later published at Odonata Central. I published my Field Key to Adult California Dragonflies (Odonata) on the Internet. State list at 108 species.
In 2001 Doug Aguillard started turning in dragonfly sightings. I shared with him my hope of someday having a dragonfly discussion group. He then started the CalOdes Yahoo! Discussion group in early 2002 and by 2006 there were at least 95 numbers for contributors/collections accessed on the California Dragonfly Distribution Excel chart.
In 2003, Tim Manolis' comprehensive
Dragonflies and Damselflies of California was
published by UCPress with Tim's own art. This year also found myself and Dave, Tim, and Andy Rehn acting as hosts for the
DSA's annual meeting. We led field trips to Bear Creek in
Colusa County, Guenoc Pond in Lake County, Pope Creek in Napa County, and Bidwell Park and Cherry Hill Campground (aka
Butterfly Valley) in Butte County. Post trips went into the Owens Valley. Although no new state records were made, 65
folks attended and Californians and California became better known in the dragonfly world:
In 2004, I updated and expanded my Common Dragonflies of California
with the new title Common Dragonflies of the Southwest, A
Beginner's Pocket Guide.
In 2005 the first annual CalOdes/DSA Dragonfly Blitz was held in Modoc County. Although no new state records were made, the camaraderie was great fun, and so was finding an Ode in the snow!!
In 2006 Azalea Creek Publishing published a dust jacket for the SW book making it more useful for Californians: Dragonflies of California and Common Dragonflies of the Southwest.
Dragonfly watching had taken off as a new pursuit and the CalOdes group was not only sharing information with each other, but actively pursuing distribution and flight season work. The rest is history and recorded on the California Sightings pages at my California Dragonfly site:
| 1977 = 102 species
1997 = 103 species #103 *Argia immunda, Kiowa Dancer - one found in Museum collection 1998 = 106 species (103 before the˙three below were added) #104 *Leucorrhinia proxima, Belted Whiteface - found for several years #105 *Aeshna canadensis, Canada Darner - found for several years #106 *Sympetrum vicinum, Autumn Meadowhawk - suspected breeder 1999= 107 species #107 *Ischnura ramburii, Rambur's Forktail - now known to breed in CA 2000 = 108 species #108 *Oplonaeschna armata, Riffle Darner - one found in Museum collection |
2001 = 108 species
2002= 108 species 2003 = 108 species 2004 = 109 species #109 *Pseudoleon superbus, Filigree Skimmer - seen only once 2005 = 109 species 2006 = 111 species #110 *Tramea calverti, Striped Saddlebags - breeding pairs; several years #111 *Erythrodiplax basifusca, Plateau Dragonlet - seen only once 2007 = 113 species #112 *Enallgama eiseni, Baja Bluet - seen only by once; some doubt about this record #113 *Rhionaeschna psilus, Turquoise-tipped Darner - seen only once |